HARVESTELLA takes place in a vibrant and colorful world where four giant crystals called Seaslight create a stable change between the four seasons. The adventure begins when abnormalities in the lake light begin to appear, creating the Quietus, a season of death that takes place between the naturally occurring four seasons. During the Quietus, crops wither and people cannot venture outside, and alarmingly, these seasons mysteriously grow longer each year.

In this life simulation RPG, players will grow and tend to harvest, use ingredients to cook and craft items, and venture into the overworld, which changes vividly depending on the season, to visit different cities, interact with townspeople, fish and more . Players can choose between different jobs, such as Fighter, Shadow Walker and Mage, and venture into dungeons with their party to discover the origin of the world and the truth behind the accident along the way.